One could argue that viral marketing is business development. The main difference is that in the former, your customers are doing the work for you, while in the latter, you've got a defined role.
I'd be careful to suggest though that this role isn't being done at all.
He introduces the list as "...that I see as important to making a new site come together."
Not "startup", "site."
I could be wrong, but Twitter doesn't need to focus on lead-gen...